Life habit: lichenized (mutualistic with algal photobionts) thallus: squamulose (squamulous) thallus: continuous, diffuse, effuse [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | green(ish) grey [th upper surface]: epruinose | pruinose [th upper surface] pruina: scarce, sparse, fine [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 1.0 (high) 4.0 [th] morphol substructures (eg areoles, squamules): discontiguous, dispersed (throughout the thallus) [th] morphol substructures (eg areoles, lobes, branches) upper surface: squamulose [th] lower surface: black(ish) [th lower surface] specific structures: present [th lower surface] rhizines, rhizoid structures: present ascomata: absent | present ascoma: perithecial, perithecioid – hymenial ascoma [mm]: (min) 0.3 ascoma: immersed, innate [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm] paraphyses/-oids: absent asci: fissitunicate – verrucarialean [asc] tholus: not thickened | thickened [asc] tholus amyloidity (iodine reaction): absent ascospores: (median) 8.0 [asp] shape: ellipsoidal | oval | clavate [asp] length [µm]: (min) 15.0 (low) 17.0(median) 18.5 (high) 22.0 (max) 24.0 [asp] width [µm]: (min) 5.0 (low) 6.0(median) 6.2 (high) 9.0 (max) 9.0 [asp] septa: absent – spore lumen unilocular, monolocular [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent secondary metabolites: absent primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.