Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: minutely squamulose squamules: loosely attached, with ± ascending, segmented, yeast-like thallus fragments with finger-like papillae; segments: 0.1-0.25 mm wide, rounded, flat or convex, upper surface: grayish green or brown, with delicate hyaline cortical hairs, anatomy: paraplectenchymatous throughout, composed of polygonal cells (c. 6-10 micro meters in diam.,) with no distinct cortices; algal cells: up to 15 micro meters diam., distributed throughout most of the thallus lower surface: similar as upper or paler Perithecia: not seen in Sonoran material; elsewhere pyriform, up to 0.4 mm high and 0.3 mm broad; wall: black, with rugose surface asci: clavate, 2-spored ascospores: muriform, brownish when old, 60-70 x 25-28micro meters Pycnidia: unknown Spot tests: all negative Secondary metabolites: none detected. Substrate and ecology: over mosses, lichens or detritus over soil or rock on dry sites World distribution: Europe (Alps, central Spain), Canary Islands, SW North America Sonoran distribution: known from a few localities in Baja California Sur and Sonora. Notes: The formation of segmented, bud-like propagules with hyaline hyphal hairs is distinctive.