Nash, T.H., Ryan, B.D., Gries, C., Bungartz, F., (eds.) 2002. Lichen Flora of the Greater Sonoran Desert Region. Vol 1.

Thallus: continuous, effuse, immersed in the substrate upper surface: pale gray or yellowish gray, glossy, smooth photobiont: absent Perithecia: hemispherical, black, semi-immersed in the thallus, c. 0.2-0.5 mm diam. involucrellum: dark brown; true exciple: 200-400 micrometer diam. ascomatal wall: black, absent below the hamathecium hamathecium: branched pseudoparaphyses, anastomosing above the asci; filaments: c. 1 micrometer wide, not inspersed with oil droplets asci: clavate, thick-walled, c. 70-100 x 17-23 micrometer, with 8, irregularly arranged ascospores ascospores: hyaline, long ellipsoid to narrowly ovoid, submuriform, 14-20 x 6-10 (15-25 x 10-15 excluding perispore - in Purvis et al. [1992]) micrometer; walls: not ornamented, without or with a c. 2-3 micrometer wide gelatinous sheath Pycnidia: black conidia: bacilliform, 8-10 x 1-1.5 mu Spot tests: all negative, UV negative Secondary metabolites: none detected. Substrate and ecology: on bark of various woody plants, mostly on Abies, Acer and Quercus World distribution: pantropical or possibly nearly cosmopolitan Sonoran distribution: common in southern Arizona, furthermore collected in Chihuahua, Sinaloa and Baja California Sur.