Global occurrence: Eurasia – Europe.  Substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: foliose (foliaceous), leaf-like; [th] upper surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | grey(ish) brown; [th upper surface]: epruinose | pruinose; [th upper surface] pruina: scarce, sparse, fine; [th marginal and upper surface] specific structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (min) 10.0; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | maculate; [th] lower surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | white(ish) | black(ish) brown; [th lower surface] specific structures: present; [th lower surface] rhizines, rhizoid structures: present; [th lower surface] rhizines, rhizinoid structures: fasciculate.  Ascomata: absent.  Secondary metabolites: present; methyl gyrophorate | phlebic acid A (28-acetoxy-22-hydroxyhopane-23-oic acid) | phlebic acid B (22-hydroxyhopane-23-oic acid) | tenuiorin | zeorin (hopane-6α,22-diol) | unknown/unidentified (tri-)terpenoid(s), (tri-)terpene(s).  Primary photobiont: present, cyanobacterial.  Secondary photobionts (eg in cephalodia): absent.