Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico) | Arctic.  Substrate: bark, cork, plant surface – trunks, branches, twigs.  Life habit: lichenized (mutualistic with algal photobionts).  Thallus: crustose (crustaceous) – endosubstratal, inconspicuous, immersed | crustose (crustaceous) – episubstratal – unspecified; continuous, diffuse, effuse | cracked, fissured, fractured, rimose | compartimentized – areolate, squamulate; [th] upper surface: grey(ish) | green(ish) grey; [th marginal and upper surface] specific structures: present; [th margin] cilia, cilioid structures: absent; [th upper surface] isidia, isidioid structures: absent; [th upper surface] soredia, soralia, soralioid structures: present; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 0.2 (high) 0.55; [th] morphol substructures (eg areoles, lobes, branches) upper surface: verrucose, warted | cracked, fissured, fractured, rimulose, subrimose.  Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 0.2 (high) 0.35 (max) 0.55; ascoma: sessile, superficial; [ascm, if apoth] disc, mazaedium: subconvex, slightly convex; [ascm, if apoth] disc, mazaedium: brown(ish) yellow (ochraceous, ochre) | red(dish) brown (if pale: orange brown) | white(ish) brown (beige); [ascm, if apoth] disc, mazaedium: epruinose | pruinose; [ascm, if apoth] disc, mazaedium pruina: scarce, sparse, fine; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: hyaline, colourless | yellow(ish) brown; [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: brown(ish) yellow (ochraceous, ochre).  Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: amyloid with acuate axial body towards the apex (= Bacidia-, Buellia-, Ramalina-types etc).  Ascospores: (median) 8.0; [asp] shape: narrowly ellipsoidal; [asp] length [µm]: (min) 7.5 (low) 9.3 (high) 10.8 (max) 13.5; [asp] width [µm]: (min) 2.5 (low) 2.8 (high) 3.3 (max) 3.5; [asp] septa: absent – spore lumen unilocular, monolocular | present, transversely septate; [asp] transversal septa: (median) 0.0 (max) 1.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent.  Secondary metabolites: present, atranorin | cryptostictic acid | isousnic acid | stictic acid | usnic acid | unknown/unidentified (tri-)terpenoid(s), (tri-)terpene(s).  Primary photobiont: present, chlorophytaceous – trebouxiaceous, chlorococcoid.  Secondary photobionts (eg in cephalodia): absent.