Global occurrence: Eurasia – Europe | Americas – North America (incl Mexico). Substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts. Life habit: lichenized (mutualistic with algal photobionts). Thallus: foliose (foliaceous), leaf-like; [th] upper surface: grey(ish) | green(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | grey(ish) brown | yellow(ish) brown; [th upper surface]: pruinose; [th upper surface] pruina: scarce, sparse, fine; [th marginal and upper surface] specific structures: present; [th margin] cilia, cilioid structures: absent; [th upper surface] isidia, isidioid structures: present; [th upper surface] soredia, soralia, soralioid structures: absent; [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 10.0 (high) 20.0; [th] morphol substructures (eg areoles, squamules): distantly discontiguous; [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | cracked, fissured, fractured, rimulose, subrimose | hairy, pilose, setose, hirsute, tomentose, pubescent, felt-like; [th] morphol substructures (eg lobes, branches): irregular; [th] lower surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) | white(ish); [th lower surface] specific structures: present; [th lower surface] rhizines, rhizoid structures: present; [th lower surface] rhizines, rhizinoid structures: unbranched, simple | fasciculate. Ascomata: absent | present; ascoma: apothecial, apothecioid – hymenial; ascoma [mm]: (low) 4.0 (high) 10.0; ascoma: sessile, superficial; [ascm, if apoth] disc, mazaedium: concave | plane, flat, flattened, expanded; [ascm, if apoth] disc, mazaedium: brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm, if apoth] disc, mazaedium: epruinose; [ascm, if apoth] margin excipular photobionts: present | sparse; [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: brown(ish) (if pale: fawn, tan; if mid: cinnamon); [ascm] paraphyses/-oids: present; [ascm] epihymenium, epithecium: brown(ish) (if pale: fawn, tan; if mid: cinnamon). Asci: lecanoralean; [asc] tholus: thickened; [asc] tholus amyloidity (iodine reaction): present; [asc] tholus amyloidity pattern: with amyloid tube, ring (= Collema-, Micarea-, Porpidia-, Psora-types etc). Ascospores: (median) 8.0; [asp] shape: acicular (needle-shaped); [asp] length [µm]: (low) 50.0 (high) 73.0; [asp] width [µm]: (low) 2.6 (high) 5.0; [asp] septa: present, transversely septate; [asp] transversal septa: (median) 3.0; [asp] pigmentation: hyaline, colourless; [asp] perispore, epispore: not apparent. Secondary metabolites: present, 15α-acetoxyhopane-22-ol (dolichorrhizin) | 7ß-acetoxyhopane-22-ol (peltidactylin) | gyrophoric acid | methyl gyrophorate | tenuiorin | zeorin (hopane-6α,22-diol) | unknown/unidentified (tri-)terpenoid(s), (tri-)terpene(s). Primary photobiont: present, cyanobacterial. Secondary photobionts (eg in cephalodia): absent.