Life habit: lichenized (mutualistic with algal photobionts) thallus: foliose (foliaceous), leaf-like [th] upper surface: brown(ish) green (olivaceous, olive green) | green(ish) brown (olive brown) [th upper surface]: epruinose [th marginal and upper surface] specific structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (min) 18.0 [th] morphol substructures (eg areoles, lobes, branches) upper surface: smooth, plane | rugose, plicate, folded, pustulate, faveolate, wrinkled | uneven, undulate [th] lower surface: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [th lower surface] specific structures: absent | present [th lower surface] rhizines, rhizoid structures: absent ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma [mm]: (min) 15.0 ascoma: sessile, superficial [ascm, if apoth] disc, mazaedium: plane, flat, flattened, expanded [ascm, if apoth] disc, mazaedium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm, if apoth] disc, mazaedium: epruinose [ascm] paraphyses/-oids: present asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): absent ascospores: (median) 8.0 [asp] length [µm]: (low) 17.0 (high) 21.0 [asp] width [µm]: (low) 5.0 (high) 6.0 [asp] septa: present [asp] septa: transversely septate [asp] transversal septa: (median) 3.0 [asp] pigmentation: pale brown [asp] perispore, epispore: not apparent conidiomata: absent | present conidia: bacilliform, bacillar [co] length [µm]: (low) 3.0 (high) 4.0 secondary metabolites: present secondary metabolites: 15α-acetoxyhopane-22-ol (dolichorrhizin) | hopane-15α,22-diol | zeorin (hopane-6α,22-diol) | unknown/unidentified compound(s) primary photobiont: present secondary photobionts (eg in cephalodia): present primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | bryophytes – mosses, liverworts.