Life habit: lichenized (mutualistic with algal photobionts) thallus: foliose (foliaceous), leaf-like [th] upper surface: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | yellow(ish) brown | red(dish) brown (if pale: orange brown) [th upper surface]: epruinose [th marginal and upper surface] specific structures: present [th margin] cilia, cilioid structures: absent [th upper surface] isidia, isidioid structures: absent [th upper surface] soredia, soralia, soralioid structures: absent [th] morphol substructures (eg areoles, lobes, branches) width [mm]: (low) 1.0 (high) 3.0 [th] morphol substructures (eg areoles, squamules): distantly discontiguous [th] morphol substructures (eg lobes, branches): subdichotomous [th] lower surface: black(ish) [th lower surface] specific structures: absent ascomata: absent | present ascoma: apothecial, apothecioid – hymenial ascoma: sessile, superficial | subpedicellate, substipitate, subpedunculate, substalked [ascm, if apoth] disc, mazaedium: brown(ish) (if pale: fawn, tan; if mid: cinnamon) [ascm, if apoth] disc, mazaedium: epruinose [ascm, if apoth] margin surface; [if perith] periostiolar area, ostiole, involucrellum: grey(ish) | brown(ish) (if pale: fawn, tan; if mid: cinnamon) | yellow(ish) brown | red(dish) brown (if pale: orange brown) [ascm, if apoth] margin excipular photobionts: present | abundant [ascm, if apoth] subhymenial layers, hypothecium; [if perith] basal excipulum: hyaline, colourless [ascm] paraphyses/-oids: present [ascm] epihymenium, epithecium: red(dish) brown (pale: orange brown) asci: lecanoralean [asc] tholus: thickened [asc] tholus amyloidity (iodine reaction): present [asc] tholus amyloidity pattern: amyloid with widening axial body towards the apex (= Lecanora-, Parmelia-, Rinodina-types etc) ascospores: (median) 8.0 [asp] septa: absent – spore lumen unilocular, monolocular [asp] pigmentation: hyaline, colourless [asp] perispore, epispore: not apparent secondary metabolites: present secondary metabolites: atranorin | physodic acid | 3-hydroxyphysodic acid | paraphysodic acid primary photobiont: present secondary photobionts (eg in cephalodia): absent primary photobiont: chlorophytaceous – trebouxiaceous, chlorococcoid.

substrate: soil, clay, humus, turf, detritus, dead leaves | wood – dead, living | bark, cork, plant surface – trunks, branches, twigs | rock – siliceous, siliciferous, acidic | rock, stones, pebbles – unspecified.