TYPE. UNITED STATES. Iowa, Fayette, on Populus, 4.XII.1895, B. Fink 194 (BM BM001106605, holotype).

Description. Life form. Facultatively lichenized fungus.

Thallus crustose, thin, continuous, white, ecorticate. Photobiont Trentepohlia or absent. Ascomata perithecia, solitary, erumpent, black, 0.2–0.4 mm diam, 0.1–0.2 mm high. Perithecial wall not carbonized, up to 100 μm thick; hamathecium hyaline, not inspersed, gel K/I+ blue; pseudoparaphyses simple, 1-1.5 μm thick. Asci cylindrical, I-, 8-spored, uniseriate; ascospores brown, fusiform with subacute ends, 3-5-septate, middle cells largest, constricted at septa, cell walls not thickened, 15-30 x 6-9 μm. Pycnidia black, of two types: 1) globose, 100-200 μm diam. with macroconidia, and 2) globose, 40-60 μm diam. with microconidia. Macroconidia 3-4-septate, bacilliform, blue-green, 12–17 x 4.5–5 μm; microconidia hyaline, simple, filiform, straight, 9–14 x 0.5 μm.

Chemistry. UV-; no substances detected by TLC.

Substrate and Habitat. Corticolous on hardwood trees.

Distribution. Temperate North America and Eurasia; in North Carolina found in the Blue Ridge ecoregion.

Literature

Aptroot., A. (1991) A Monograph of the Pyrenulaceae (Excluding Anthracothecium and Pyrenula) and the Requienellaceae, with Notes on the Pleomassariaceae, the Trypetheliaceae and Mycomicrothelia (Lichenized and Non-lichenized Ascomycetes). Bibliotheca Lichenologica 44: 1-178. 

Ezhkin, A.K. & F. Schumm. (2018) New and noteworthy records of lichens and allied fungi from Sakhalin Island, Russian Far East, II. Folia Cryptogamica Estonica 55: 45-50.

Harris, R.C. & D. Ladd. (2005) Preliminary Draft: Ozark Lichens, enumerating the lichens of the Ozark Highlands of Arkansas, Illinois, Kansas, Missouri, and Oklahoma. Published by the authors. 249 pp.